Trypeticus fagi (Lewis, 1884)


Tryponaeus fagi Lewis, 1884: 138 [Japan, no further locality].

Trypeticus fagi: Lewis, 1905d: 7; 1912: 251 [male and female genitalia illustrated].

Japanese name: Tsutsu-emma-mushi.

Trypeticus fagi: Ôhara, 1994: 267.


Original description. "Cylindricus, niger, nitidus, undique punctatus; antennis pedibusque rufo-piceis. L. 4 mill.

Male. Frontis lateribus basisque margine elevatis; pronoto antice rectuso, medio subelevato.

Female. Fronte plana, in medio depressa; pronoto antice convexo. "


Redescription (Ohara, 1994)

Male: Body length, PPL, male, 3.11 - 3.50 (3.36 ± 0.04, n=10) mm, female, 3.11 - 3.60 mm (3.42 ± 0.06, n=7), PEL, male, 2.84 - 3.11 mm (2.99 ± 0.03, n=10), female, 2.94 - 3.14 mm (3.07 ± 0.02, n=7). Width, male, 1.31 - 1.34 mm (1.31 ± 0.001, n=10), female, 1.24 - 1.31 mm (1.28 ± 0.01, n=7). Biometric data are given in Table 54. Body cylindrical, moderately stout. Cuticle shining, black; tibiae, tarsi, and antennae dark rufopiceous.

Head without frontal stria (Fig. 159C, D); surface slightly depressed on median area, and sparsely clothed with fine punctures which are separated by two to five times their diameter and become coarser posteriorly; surface among the punctures extremely finely covered with alutaceous ground sculpture.

Anterior margin of pronotum shallowly emarginate on median four-sixths; sides parallel (Fig. 158) and straight; marginal stria complete and carinate laterally, and absent behind head on anterior margin; surface of disk densely and coarsely punctate, but not on longitudinal mid line, the punctures becoming denser on apical one-fourth and on area along lateral margin; surface among coarse punctures usually impunctate and shining, but sometimes covered with fine punctures, which are one-third as large as the coarse punctures.

Epipleura broad, not concave. Marginal epipleura and elytral stria absent. Narrow band along epipleural margin impunctate, but extremely finely striate. Surface of elytra irregularly scattered with coarse punctures which are somewhat smaller than those of the pronotum and usually separated by two to five times their diameter, the punctures becoming coarser on basal margin and denser along the sutural line and lateral area along epipleura; a small oblong area on humeral region impunctate; extreme apex of elytra impunctate.

Disk of propygidium (Fig. 159) evenly covered with round and shallow punctures, which are a little coarser than the elytral ones and separated by about their diameter, and with other fine punctures intermingled. Punctation of pygidium similar to that of propygidium, but much sparser, the punctures becoming finer apically; tip of pygidium clearly acute.

Prosternal process longitudinal quadrangular; anterior margin straight; marginal stria of the process nearly complete and somewhat carinate; disk densely covered with coarse and shallow punctures and intermingled with fine punctures among the coarse ones; surface among these punctures finely clothed with alutaceous ground sculpture.

Median third of anterior margin of mesosternum nearly straight; lateral third inwardly curved, fitting to posterior margin of procoxa; lateral sides nearly parallel and straight on basal half and divergent apically on apical half; each lateral side marginate and deeply carinate; punctation of disk similar to that of prosternum. Meso-metasternal suture impressed and obtuse-angulate (about 120°) at middle. Metasternum clearly depressed on longitudinal mid line; lateral metasternal striae slightly divergent posteriorly and impressed on basal two-thirds. Intercoxal disk of metasternum sparsely covered with coarse, somewhat longitudinally oblong and shallow punctures which are separated by their own diameter to thrice the dimeter; surface among coarse punctures sparsely and finely punctate except on a broad band along longitudinal mid line. Lateral disk of metasternum sparsely and coarsely punctate, the punctures much sparser than those of the intercoxal disk. Metepisternum with two rows of dense and coarse punctures.

Table 54. Biometeric data of Trypeticus fagi (Lewis).

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Male Female

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PW 1.21-1.31 (1.27± 0.01) 11 1.24-1.31 (1.27±0.01) 7

PL 1.44-1.57 (1.49± 0.01) 11 1.47-1.64 (1.57±0.02) 7

EL 1.31-1.54 (1.46± 0.02) 11 1.34-1.50 (1.43±0.02) 7

EW 1.31-1.34 (1.31±0.001) 10 1.24-1.31 (1.28±0.01) 7

ProW 0.72-0.85 (0.79± 0.01) 11 0.69-0.82 (0.75±0.02) 7

ProL 0.29-0.36 (0.33± 0.01) 11 0.26-0.33 (0.30±0.01) 7

PyL 0.49-0.59 (0.52± 0.01) 11 0.49-0.65 (0.57±0.02) 7

PTL 0.52-0.65 (0.62± 0.01) 11 0.52-0.62 (0.58±0.01) 7

MSTL 0.49-0.65 (0.59± 0.02) 11 0.52-0.65 (0.58±0.02) 7

MTTL 0.59-0.69 (0.63± 0.01) 11 0.59-0.65 (0.62±0.01) 7

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Intercoxal disk of 1st abdominal sternum densely covered with coarse punctures which are a little smaller than those of the metasternal intercoxal disk, the punctures becoming sparser on basal half.

Protibia slender, with 5 large denticles on outer margin. Mesotibia with 5 denticles on outer margin.

Female. In general appearance similar to the male except in the following characters: head (Fig. 159A, B) with clear frontal marginal stria which is strongly carinate; pronotum distinctly with short costa on apical fourth on longitudinal mid line, and area around the costa densely clothed with longitudinal short rugae and fine alutaceous ground microsculpture where it is lusterless; pronotum evenly punctate wholly, without impunctate area on longitudinal mid line; punctation of pygidium much denser than that of the male; tip of pygidium round, not acute; prosternal process without marginal stria.

Male genitalia as shown in Fig. 160.


Specimens examined. (Ohara, 1994)

[Honshû] <Aomori-ken> 1Åä, no date, A. Abe leg.; 1Åâ, Oirase, Nishigun, Fukaura-machi, 23/v/1976, A. Abe leg.; 1Åâ, Yachi Spa., Towada, 26/vii/1952, Y. Kurosawa leg. <Fukushima-ken> 1Åâ, Yunohana, Minami-Aizu, 20/vii/1963, K. Tazoe leg.; 1Åä, Nagasaka, Iide-san, 14/vi/1970, K. Tazoe leg.; 1Åâ, Hinoemata, 30/v/1982, H. Makihara leg.; 1Åä, Shin-machi, Tateiwa, Minami-Aizu, 18/vi/1947, Y. Kurosawa leg. <Saitama-ken> 1Åâ, Yanagi-koya, Chichibu, 22/vii/1933, no collector's name. <Gifu-ken> 1Åâ, Idani, Kawai, Hida, 5/vi/1955, H. Torigai leg.; 2ÅâÅâ, Ditto, 1/vi/1957, T. Nakane leg. <Mie-ken> 1Åâ, Mie Univ. Forest., Ichishi-gun, 11/vi/1956, H. Ichihashi leg.; 1Åä, Ditto, 12/vi/1956, H. Ichihashi leg. <Kyôto-fu> 1Åä, Ashiu, vi/1951, A. N leg.

[Shikoku] <Tokushima-ken> 1Åâ, Ashû, 27/vii/1940, M. Tokunaga leg. (Host: Aesculus turbinata Blume). <Ehime-ken> 1Åä, Omogo, 11/vii/1956, Z. Naruse et M. Satô leg.

[?] 1Åâ1Åä, Sakasimo, Hora, 13/vi/1954, H. Torigai leg.


Specimens examined [additional records]. (Ohara, 1999a).

[Honshu] Aomori-ken: Towada, Hiraka-machi (4 exs., 25, 26/vi/1964; 1 ex., 9/viii/1967, APM), K. Shimoyama; Juniko, Iwasaki-mura (2 exs., 21/vi/1964, APM), K. Shimoyama; Mt. Kushiishiyama, Ajigasawa-machi (1 ex., 7/vi/1987, APM), A. Abe; (1 ex., 30/vii/1987, APM), Y. Kikuchi; Jôgakura, Mts. Hakkôda (2 exs., 22/vi/1986, APM), A. Abe; Ôkawa, Nishimeya-mura (1 ex., 25/vii/1987, APM), A. Abe; Ajigasawa-machi (1 ex., 5/viii/1986, APM), A. Abe; Akaishigawa, Ajigasawa-machi (1 ex., 27/vii/1986, APM), K. Shimoyama; Mt. Ranganmori, Ajigasawa-machi (1 ex., 6/viii/1986, APM), S. Yamauchi.


Distribution. Japan (Honshu, Shikoku, Kyushu, Izu Isles, Yaku-shima Is.).



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