Atholus depistor (Marseul, 1873)
Hister depistor Marseul, 1873, 176.
Hister depistor Marseul, 1873, 224 [Japan: Nangasaki (Kiu-siu = Kyushu)].
Hister (Peranus) depistor: Bickhardt, 1910, 53 [catalogued]; 1917, 192 [catalogued]; Müller, 1937, 130 [Japan; Mongol]; Kamiya & Takagi, 1938, 31 [listed]; Ôsawa & Nakane, 1951, 6 [noted; figured].
Peranus depistor: Lewis, 1906, 402; 1915, 55 [Formosa]; Nakane, 1963, 70.
Atholus (Atholus) depistor: Kryzhanovskij & Reichardt, 1976, 386; Nakane, 1981, 10 [listed]; Hisamatsu & Kusui, 1984, 17 [key; noted]; Hisamatsu, 1985, 228, pl. 41, f. 17 [noted; key; photo].
Atholus depistor: Ôhara, 1993, 137.
Peranus depistor: Lewis, 1915, 55 [Horisha]; Mazur, 1970, 59 [Korea].
Hister (Peranus) depistor: Miwa, 1931, 57 [Horisha]; Kamiya and Takagi, 1938, 31; Reichardt and Kryzhanovskij, 1964, 174 [Korea (Seoul)].
Atholus depistor: Ohara, 1992, 176 [Taiwan: Kenting Park]; Ohara, 1999: 32.
Atholus (Atholus) depistor: ESK & KSAE, 1994, 137 [Korea].
Ohara (1993)
Description. Male and female. Body length, PPL: male, 4.43 - 5.47 mm, female, 4.66 - 6.19 mm, PEL: male, 3.83 - 4.43 mm, female, 3.95 - 4.90 mm. Width: male, 2.86 - 3.81 mm, female, 3.33 - 4.24 mm. Biometric data are given in Table 3. Body oblong-oval, convex, black and shining; antennae, tibiae, and tarsi dark brown.
Frontal stria of head (Fig. 6 A) complete, carinate, densely crenate and inwardly angulate at middle. Disk wholly covered with reticulous ground sculpture, and sparsely covered with fine punctures which are separated from one another by about three times their diameter. Labrum transversely oblong. Dorsal surface of mandible with a strong carina along outer margin.
Pronotal sides (Fig. 6 B) regularly arcuate, but on the apical half strongly convergent anteriorly. Apical angles acute. Marginal pronotal stria present on apical half laterally and broadly interrupted anteriorly behind head. Lateral pronotal stria strongly impressed and densely crenate, the lateral portion shortened on basal fourth and rather widely distant from the pronotal lateral margin. Area within the antero-lateral angle of lateral stria deeply excavated and densely and largely punctate. Disk wholly covered with reticulous ground sculpture, and sparsely scattered with fine punctures. Ante-scutellar area with a short longitudinal puncture.
Epipleura of elytra deeply and longitudinally excavated, and densely and coarsely punctate. Marginal epipleural stria clearly impressed on apical two-thirds along epipleural margin. Marginal elytral stria deeply impressed and complete. Lateral margin of elytron strongly carinate. External subhumeral stria (Fig. 6 B) present briefly on median portion, as an arc. Internal subhumeral stria variable, shortly present medially. These subhumeral striae often fused with each other, the apical end of the external stria and the basal end of the internal one united into a continuous stria, which is half to two-thirds as long as elytra. Oblique humeral stria lightly impressed on basal third. First to 5th dorsal striae complete and sparsely and coarsely crenate, the 5th outwardly arcuate. Sutural stria present on apical two-thirds or less. Disk of elytra wholly and coarsely covered with reticulous sculpture.
Propygidium (Fig. 9 C) evenly and coarsely covered with round and deep punctures which are separated from one another by half to one their diameter, and sparsely intermingled with fine punctures; interspace among punctures wholly covered with microscopic alutaceous ground sculptures. Punctation of pygidium (Fig. 9 H) similar to that of propygidium, but the punctures are much denser and become finer apically.
Anterior margin of prosternal lobe (Fig. 6 C) narrowly truncate medially; marginal stria deeply impressed, carinate and complete, the lateral portion rather distant from lateral margin; disk sparsely and finely punctate medially and densely covered with coarse punctures laterally. Prosternal keel narrow, the anterior half descending; carinal stria absent; disk sparsely clothed with fine punctures which are separated by about three times their diameter, the punctures becoming coarser on sides of apical half. Descending lateral stria deeply impressed, carinate and complete.
Anterior margin of mesosternum (Fig. 6 D) outwardly arcuate, but the median portion is slightly and narrowly emarginate; marginal stria strongly carinate, sparsely and coarsely crenate, and complete; a short oblique stria present between the marginal stria and the antero-lateral angle on each side; disk evenly covered with fine punctures which are separated by about four times their diameter. Meso-metasternal suture complete, obtusely angulate at middle. Lateral stria of metasternum deeply impressed, carinate, obliquely and posteriorly extending, and beginning from lateral fourth of meso-metasternal suture. Post-mesocoxal stria carinate and complete, somewhat broadly distant from posterior margin of mesocoxa, the basal end united with apical end of marginal stria of mesosternum. Punctation of intercoxal disk of metasternum similar to that of mesosternum. Lateral metasternal disk densely covered with large, shallow, round and setiferous punctures, the punctures finer along the lateral stria.
Intercoxal disk of 1st abdominal sternum (Fig. 6 D) sparsely and finely punctate, the punctures separated by two to three times their diameter; 1st abdominal sternum deeply striate on each side, the stria carinate and complete.
Protibia (Fig. 6 E & F) with 5 denticles on outer lateral margin, the basal one very small and the apical two appressed together on apical angle, and with one denticle on apical margin. Profemur (Fig. 6 G) with femoral stria nearly complete and most distant from the posterior margin at middle. Mesotibia and metatibia each with 8 long and 9 to 10 short spines on outer margin.
Male genitalia as shown in Fig. 7.
Female genitalia as shown in Fig. 6 H.
Remarks. This species is similar to Atholus duodecimstriatus quatuordecimstriatus, but is separated from the latter by the characters given in the key and by having an excavation within the anterior angle of pronotal lateral stria.
Table 3. Biometric data for Atholus depistor (Marseul).
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Part measured Male Female
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APW 1.09-1.55 (1.38+0.02) 20 1.33-1.67 (1.49+0.02) 20
PPW 2.62-3.38 (3.11+0.04) 20 2.88-3.76 (3.37+0.06) 20
PL 1.29-1.67 (1.51+0.02) 20 1.43-1.86 (1.64+0.03) 20
EL 1.95-2.38 (2.24+0.03) 20 2.24-2.81 (2.46+0.04) 20
EW 2.86-3.81 (3.43+0.05) 20 3.33-4.24 (3.75+0.06) 20
ProW 1.90-2.28 (2.07+0.02) 20 2.00-2.52 (2.29+0.04) 20
ProL 0.86-1.00 (0.92+0.01) 19 0.86-1.14 (0.99+0.01) 20
PyL 0.81-1.09 (0.94+0.02) 20 0.86-1.29 (1.07+0.03) 20
PTL 0.95-1.24 (1.12+0.02) 19 0.95-1.38 (1.18+0.02) 20
MSTL 0.86-1.24 (1.04+0.02) 19 0.95-1.29 (1.09+0.02) 20
MTTL 1.05-1.43 (1.28+0.02) 19 1.19-1.57 (1.37+0.02) 20
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Specimens examined. 19ÅâÅâ, 20ÅäÅä and 32 exs.
[Hokkaido] 1Åâ, Sôunkyô, Daisetsu, 28-VII-1985, M. Ôhara leg.; 8ÅâÅâ, 5ÅäÅä, 4 exs., Jôzankei, 20-VI, 27-VIII, 14-IX, 13-X-1985, M. Ôhara leg.; 2 exs., Toyotaki, Sapporo, 13-X-1985, 23-VIII-1987, M. Ôhara leg.; 2ÅäÅä, Hitsujigaoka, Sapporo, 20-VIII-1985, M. Ôhara leg.; 1Åâ, Iwanai, 9, 10-VII-1990, M. Ôhara leg.; 1 ex., Ômori-hama, Hakodate, 15-VIII-1952, K. Homma leg. (EIHU). <Okushiri Is.> 1Åä, 28-VI-1956, K. Umeya leg. (EIHU).
[Honshu] <Fukushima-ken> 1 ex., Wakamatsu, 16-VIII-1948, K. Nagayama leg. (NSMT). <Chiba-ken> 1 ex., Ôami, 22-VI-1984, M. Ôhara leg. <Saitama-ken> 1 ex., Adachi, Arakawa, 31-V-1970, M. Ishida leg. <Tokyo-to> 1 ex., Fuchû, 11-VI-1968, M. Ishida leg. <Kanagawa-ken> 2ÅâÅâ, 2ÅäÅä, 4 exs., Akuwa-chô, Yokohama, 3-VI-1983, K. Wada leg.; 1Åâ, Matsudayama, Tanzawa, 14-IX-1975, M. Ishida leg.; 1Åâ, 2ÅäÅä, 2 exs., Fujisawa, 14-VI-1984, I. Maehara leg. <Niigata-ken> 1 ex., Senami, northern Echigo, 21-VI-1985, K. Baba leg. <Nagano-ken> 1 Åä, Noziri, 24-VII-1930, T. Ishida leg. (NA); 1Åä, ditto, 21-VIII-1943, T. Nakane leg. (NA). <Mie-ken> 1Åä, 1 ex., Yunoyama, 6-VIII-1973, M. Tanaka leg.
[Kyushu] <Fukuoka-ken> 1 ex., Fukuoka, 16-VI-1948 Y. Abitsu leg. (NA); 1 ex., Tashiro, 23-VI-1948, Y. Miyake leg. (NA); 1 ex., Mt. Asaka, 23-VIII-1946, F. Takahashi leg. (NA).; 1 ex., Mt. Adachi, 9-VI-1969, S. Nakao leg. (NA). <Kumamoto-ken> 1Åâ, Haruyama, Takamori, Aso-gun, 6-VI-1986, E. Matsui leg.
[Nansei Isls.] <Yakushima Is.> 1 ex., 2-IV-1985, M. Kiuchi leg.; 1 Åä, Kurio, 5-X-1982, M. Ôhara leg. <Okinawa-hontô Is.> 1 ex., Ôgimi, 18-V-1984, M. Shimabukuro leg. <Miyako Is.> 1 ex., 1-VI-1969, Y. Kusui leg. <Zamami Is.> 1Åâ, 3ÅäÅä, 27-VII-1987, Y. Kusui leg. <Iriomote Is.> 2 exs., Ôtomi, 16-III-1985, K. Nakamine leg. <Ishigaki Is.> 1 ex., Isobe, 29-III-1984, Yoshida leg.; 1 ex., Kabira, 5-VI-1969, Y. Kusui leg.; 3ÅâÅâ, 1Åä, Hirakubozaki, 24-VIII-1987, H. Tanaka leg.
[Ogasawara Isls.] 1 ex., Komagari, Chichi-jima, 8-IV-1973, Y. Kusui leg.
[Taiwan] 1 ex., Kenting Park, Pingtung, 11-XI-1976, M. Kiuchi leg.
[Continental China] 1 ex., Sinkyo (= Changchun), Manchukuo, 14-VII-2599(=1939), H. Tokuhiro leg. (NSMT).
Jeonranam-do: Cheongup, 24/vi/1966 (1 ex., W. H. Paik). Cheju-do: Ha-gwi,
4/viii/1993 (2 exs., J.-C. Paik).
Japan: [Honshu] Chiba-ken: Odake (#2623), Sakura-shi (1 ex., 12/ix/1997, CBM), M. Maruyama and M. Sano; Iizuka (#3439), Sakura-shi (1 ex., 29/ix2/x/1997, carrion baited trap, CBM), M. Maruyama et M. Sano. Kanagawa-ken: Jyusan-toge, Nagaura, Yokosuka (1 ex., 2/iv/1963, YCM), T. Hamaguchi.
[Nansei Isles] Iriomote-jima: Uehara (15 exs., 15/ix/1993, collected from cow dung), M. Ohara.
Specimens examined (Ohara, 1999b).
Taiwan: [Proper] Nantou Hsien: Nanshanchi, Puli (3 exs., 13/iv/1986), M. Ohara; Puli (1 ex., 8/x/1976), M. Kiuchi; Chitou (4 exs., 5/viii/1977), native collector. Hualien Hsien: Jui-sui (1 ex., 15/xi/1976), M. Kiuchi. Pingtung Hsien: Kenting park (1 ex., 11/xi/1976) M. Kiuchi; (1 ex., 4/iv/1976), K. Tazoe. ?: Taiwan Botanical Garden (2 exs., 2/viii/1968), T. Kosaka.