Saprinus planiusculus Motschulsky, 1849: 97 [Kirgiz SSR].
Saprinus cuspidatus Ihssen, 1949: 183, synonymized by Kryzhanovskij, 1972: 435.
Saprinus nitidulus: Marseul, 1873: 221 [Japan; Shimabara (Kiu-Siu) et Hiogo (Niphon)].
Japanese name: Dôgane-emma-mushi.
Saprinus (Saprinus) planiusculus: Ôhara, 1994, 232.
Saprinus planiusculus Motschulsky, 1849, 97.
Saprinus cuspidatus Ihssen, 1949, 183; Mazur, 1970, 57 [Korea].
Saprinus (Saprinus) planiusculus: Ohara, 1994, 232.
Japanese name: Dôgane-emma-mushi.
Description. Body length, PPL, male, 3.95 - 5.66 mm (4.72 ± 0.08, n=20), female, 4.09 - 6.33 mm (5.30 ± 0.12, n=20), PEL, male, 3.52 - 4.71 mm (4.20 ± 0.06, n=20), female, 3.81 - 5.09 mm (4.47 ± 0.08, n=20). Width, male, 3.14 - 4.19 mm (3.64 ± 0.05, n=20), female, 3.28 - 4.57 mm (3.94 ± 0.07, n=20). Biometric data are given in Table 46. Body broadly oval and strongly convex. Cuticle very strongly shining with brassy lustre, black; tarsi and funicle of antennae dark rufopiceous.
Frontal stria of head weakly carinate and complete. Supraorbital stria well impressed and complete. Disk of front densely covered with moderate punctures, which become sparser on the basal half; interspace among these punctures usually smooth, occasionally clothed with fine punctures. Epistoma medio-apically more densely punctate than the disk. Labrum deeply depressed medially.
Pronotal sides (Fig. 132) evenly arcuate and convergent on basal five-sixths,
thence strongly convergent to apices. Apical angles obtusely angulate. Marginal
stria carinate and complete. Disk smooth, but densely, coarsely and deeply
punctate along the sides except on basal fourth, sparsely clothed with fine
punctures in a narrow band outside the coarsely punctate area; base with
two or three irregular rows of large punctures except on lateral seventh
and a narrow antiscutellar area; elsewhere extremely finely, sparsely punctate;
area behind eyes feebly depressed.
Epipleura of elytra sparsely with moderate punctures, which become denser on the apical third. Marginal epipleural stria complete and finely impressed. Marginal elytral stria lightly carinate and complete, its apical end extending along the posterior margin of elytra to medio-apical angles of elytra and then bending basally and running for a short distance. External subhumeral stria confined basally. Internal subhumeral stria present on median third, with coarse punctures, and sometimes its basal end united with the apical end of oblique humeral stria. Oblique humeral stria (Fig. 132) distinctly impressed on basal third. First, 2nd and 4th dorsal striae present on about basal half, and coarsely and sparsely crenate; 2nd usually a little longer than the 1st. Third dorsal stria short, present on basal fifth, and coarsely crenate. Fifth dorsal stria wanting. Sutural stria shortly represented by a row of four to five coarse punctures medially. Surface of apical half or third of elytra covered with coarse and oval punctures, which are finer than those in an area along the pronotal sides, and usually separated by their own diameter to twice the diameter, but sometimes sparser; the punctate area usually expanding basally onto sutural area and also interval between 2nd and 4th dorsal striae; interval between 1st and 2nd dorsal striae usually with several longitudinal and short lines; surface of basal half smooth, sparsely clothed with microscopic punctures; extreme apex of elytra (around apical stria) sparsely and finely punctate; interspace among all discal punctures of elytra smooth, occasionally clothed with microscopic punctures.
Propygidium densely covered with coarse punctures, which are a little coarser than the elytral coarse punctures and become finer on basal half; interspace among the punctures occasionally and finely punctate and with lightly impressed, fine and irregularly alutaceous microsculptures. Pygidium densely covered with punctures, which are slightly coarser than those of the propygidium and progressively become finer on the apical third and nearly absent at the apex.
Anterior margin of median portion of prosternum straight. Ventral disk of keel even, sparsely clothed with microscopic punctures, which are coarser laterally. Anterior marginal stria of keel deeply impressed, carinate, and complete. Carinal stria carinate, complete, divergent anteriorly and posteriorly, and nearly straight on apical third. Descending lateral stria strongly carinate, its apical end attaining to apical third of carinal stria.
Anterior margin of mesosternum broadly and feebly emarginate medially. Marginal stria complete, and strongly carinate. Disk sparsely clothed with microscopic punctures, which become coarser laterally. Meso-metasternal suture lightly impressed, and accompanied with a strongly crenate line. Intercoxal disk of metasternum feebly convex, sparsely clothed with microscopic punctures, which are coarser along lateral metasternal stria and in a transverse apical band. Lateral metasternal stria strongly carinate, extending obliquely and posteriorly, and present usually on basal half. Lateral metasternal disk densely covered with round, shallow, large and setiferous punctures, which become smaller on the apical third, with fine punctures intermingled. Metepisternum more densely punctate than those of lateral disk of metasternum, the punctures becoming sparser on apical half.
Intercoxal disk of 1st abdominal sternum with strongly carinate lateral stria on basal three-fourths: sparsely and microscopically punctate on median area; lateral area along lateral stria sparsely and coarsely punctate, with fine punctures intermingled.
Protibia with 13 spines on outer margin, the apical two and basal three small.
Male genitalia as shown in Fig. 135.
Ohara (1994: Fig. 135)
Table 46. Biometeric data of Saprinus planiusculus Motschulsky.
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Male Female
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APW 1.19-1.43 (1.32±0.01) 20 1.24-1.57 (1.42±0.02) 20
PPW 2.81-3.81 (3.36±0.05) 20 3.05-4.00 (3.61±0.06) 20
PL 1.33-1.86 (1.58±0.03) 20 1.38-1.86 (1.68±0.03) 20
EL 2.28-2.86 (2.57±0.03) 20 2.24-3.19 (2.70±0.06) 20
EW 3.14-4.19 (3.64±0.05) 20 3.28-4.57 (3.94±0.07) 20
ProW 2.05-3.05 (2.29±0.04) 20 2.00-3.05 (2.42±0.05) 20
ProL 0.48-0.71 (0.58±0.02) 20 0.43-0.62 (0.54±0.01) 20
PyL 1.29-1.81 (1.54±0.03) 20 1.52-2.14 (1.81±0.03) 20
PTL 1.14-1.52 (1.34±0.02) 20 1.14-1.67 (1.37±0.03) 20
MSTL 1.29-1.76 (1.46±0.02) 20 1.19-1.67 (1.42±0.03) 20
MTTL 1.48-2.00 (1.68±0.03) 20 1.38-1.95 (1.68±0.04) 20
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Remarks. Saprinus planiusculus resembles Saprinus niponicus, with which it is to a great extent sympartic; however, it can be distinguished from the latter by the shortened 3rd dorsal elytral stria and the shape of the 8th sternum of the male genitalia.
This species occurs on dead animal bodies.
Specimens examined. [Hokkaidô] 1 ex., Shibecha, Kushiro, 16/vi/1955, K. Ijima leg. (IJ); 1 ex., Kamui-kotan, Asahikawa, 20/vii/1987, M. Ôhara leg.; 1Åâ, Ombetsu, 25/vi/1985, M. Ôhara leg.; 1 ex., Otaru, Shiribeshi, 1/vii/1943, T. Hasegawa leg.; 3 exs., Ishikari-hama, 7/viii/1978, T. Fujisawa leg.; 2 exs., Zenibako, 27/vi/1985, 19/vi/1985, M. Ôhara leg.; 5 exs., Sapporo, 19, 27/vi/1985, 9/vii/1987, M. Ôhara leg.; 1 ex., Maru-yama, Sapporo, 23/vi/1979, N. Nishikawa leg.; 1 ex., Ditto, 2/vi/1951, T. Kumata leg. (EIHU); 1 ex., Jôzankei , Sapporo, 8/vii/1907, S. Matsumura leg. (EIHU); 1 ex., Misumai, Sapporo, 27/vi/1990, M. Ôhara leg.; 18 exs., Ditto, 12-VII, 9, 10 & 18/vii/1991, K. Sayama leg.; 13 exs., Nopporo, 12/vi/1986, M. Ôhara leg.; 9 exs., Kamayausu, Sapporo, 20/vii/1987, M. Hinakura leg.; 1Åâ1Åä, Hakodate, 19/vi/1954, T. Nakane leg. (Saprinus cuspidatus, Dahlgren det.) (NA); 1 ex., Hakodate, 17/vi/1954, T. Nakane leg. (NA); 1 ex., Mt. Hakodate, 26/viii/1951, K. Honma leg. (NA); 2 exs., Yagishiri Is., 21/vi/1985, Y. Kusui leg.
[Honshû] <Iwate-ken> 1 ex., Yashikidai, 19/vii/1954, T. Kumata leg. (EIHU). <Yamagata-ken> 12 exs., Yonezawa, 16/v/1944, Y. Kurosawa leg. (NSMT). <Fukushima-ken> 1 ex., Wakamatsu, 29/vi/1049, T. Nakane leg. (NA); 2 exs., Mt. Komagatake, southern Aizu, 10/viii/1949, K. Nagayama leg. (NSMT); 2 exs., Wakamatsu, 14/v/1947, Y. Kurosawa leg. (NSMT); 1 ex., Fukushima-shi, 9/vii/1965, K. Tazoe leg. <Chiba-ken> 2 exs., Cape Futtu, 2/vii/1961, M. Yamazaki leg. (NA); 1 ex., Ôami, 19/vi/1984, M. Ôhara leg.; 2 exs., Ichikawa, 27/iv/1958, M. Suwa leg. <Tôkyô-to> 2 exs., Ogikubo, 16/vi/1963, 21/iv/1968, M. Ishida leg.; 1 ex., Kumegawa, Higashimurayama, 26/iv/1964, M. Ishida leg.; 1 ex., Setagaya, 18/ix/1949, Ôhira leg. (NA); 1 ex., Takamizu-yama, 2/vi/1935, T. Nakane leg. (NA); 1Åä, 1 ex., Hongô, Musashi, v/1947, T. Nakane leg. (Saprinus cuspidatus, Dahlgren det.) (NA); 2 exs., Aoyama, 26/vii/1943, 25/v/1948, T. Nakane leg. (NA); 1 ex., Komazawa, 23/v/2603 (= 1943), Watanabe leg. (NSMT); 1 ex., Kinuta, 16/v/1965, Y. Kurosawa leg. (NSMT); 1 ex., Ueno-kôen, 1/vii/1934, H. Uchida leg. (NSMT); 1 ex., Hatagaya, 11/v/1924, no collector's name. <Yamanashi-ken> 1 ex., Masutomi, 12/vi/2603, no collector's name. (NSMT); 1 ex., Sagashiro Spa., 28-30/vii/1961, S. Nomura leg. (NSMT). <Niigata-ken> 1 ex., Niigata-shi, 26/vi/1933, K. Baba leg. <Nagano-ken> 1 ex, Fukushima, Kiso, 11-12/vi/1940, T. Nakane leg. (Saprinus cuspidatus, Dahlgren det.) (NA); 1 ex., Kami-kôchi, Shinano, 21-28/vii/1947, S. Ôsawa leg. (NA); 3 exs., Sunama, Nojiri, 21/viii/1943, T. Nakane leg. (NA); 1 ex., Gaijin-mura, Nojiri, 19/viii/1941, T. Nakane leg. (NA). <Aichi-ken> 3 exs., Obata, Owari, 15/vi/1941, 30/v/1947, S. Ôsawa leg. (NA); 1 ex., Miya, Mikawa, 1-2/ix/1947, S. Ôsawa leg. (NA); 1 ex., Shiadeki-machi, Nagoya, 1/vi/1941, S. Ôsawa leg. (NA). <Gifu-ken> 3 exs., Hirayu, 24/vii/1946, M. Hayashi leg. <Hyôgo-ken> 1 ex., Motoyama-mura, near Kôbe, 14/v/1938, K. Sakaguchi leg. (NA); 1 ex., Kôbe, 22/vi/1930, no collector's name. (BSM). <Tottori-ken> 2 exs., Daisen, 27/v/1935, M. Hamano leg. (NSMT).
[Kyûshû] <Fukuoka-ken> 1 ex., Hakozaki, 24/vi/1946, F. Nakasuji leg. (NA).
[Hokkaido] Rishiri Is.: Numaura (1 ex., 29/vii/1994), M. Ohara. Oshima-Oshima Is.: Yamase-domari (4 exs., 25/vi/1994, NIS), H. Nishijima; Torikarasu (1 ex., 23/vi/1994, NIS), H. Nishijima. Okushiri Is.: Miyatsu (1 female, 3/ix/1986), Y. Kusui; Aonae (1 male & 1 female, 28/vi/1986), Y. Kusui. Proper: Miyama, Kushiro-chô (15 exs., 25/vii/1997, IJ), K. Ijima; Toyohama, Shimamaki (1 ex., 313/vi/1996, SA), K. Sasaki; Yahata, Ishikari (1 ex., 22/v/1994, SA), K. Sasaki; Makomanai, Sapporo (1 ex., 24/viii/1995, SA), K. Sasaki.
[Honshu] Aomori-ken: Chikagawa, Mutsu-shi (1 ex., 16/vi; 3 exs., 2/vii; 8 exs., 3/vii; 4 exs., 6/vii; 1 ex., 9/vii/1982; APM), K. Shimoyama & A. Satoh; (1 ex., 2/vii/1982, APM), A. Satoh; Araya, Hiraka-machi (1 ex., 9/i/1971, APM), A. Satoh; Towada, Hiraka-machi (1 ex., 1/xi/1981, APM), K. Shimoyama; Nakajima, Jûsanko, Shiura-mura (13 exs., 17/vi/1984, APM), no collector's name; Iizume, Goshogawara-shi (1 ex., 2/x/1972, APM), A. Abe; Oirasegawa, Fukaura-machi (1 ex., 16/viii/1975, APM), K. Shimoyama; Mt. Rangannomori, Ajigasawa-machi (1 ex., 13/vi/1971, APM), A. Abe. Chiba-ken: Aoba, Chiba-shi (1 ex., 13/ix/1986, CBM-Z-53808), H. Yamazaki. Kanagawa-ken: Hiratsuka (1 ex., 30/viii/1963), Y. Kurosa. Nagano-ken: Sugadaira (1 ex., 31/viii/1969, IMA), M. Fujioka.
