Saprinus optabilis Marseul, 1855: 438; Bickhardt, 1913: 177
[Kosempo]; Lewis, 1915: 56 [Koshun]; Miwa, 1931: 51 [Kôshun];
Dahlgren, 1968: 261 [figures of genitalia]; Mazur and Ôhara,
2003: 8 [Thailand].
Saprinus dives Lewis, 1911: 88, synonymized by Dahlgren, 1969:
266.
Saprinus (Saprinus) optabilis: Mazur, 1997: 226.
Ohara (2004)
Redescription. Body length*, PPL, male, 3.92 5.14 mm
(4.54 ± 0.13, n=12), female, 4.02 5.64 mm (4.95 ±
0.12, n=12), PEL, male, 3.48 4.56 mm (4.11 ± 0.10,
n=12), female, 3.72 4.80 mm (4.48 ± 0.09, n=12).
Width, male, 2.94 3.77 mm (3.38 ± 0.08, n=12), female,
3.09 4.08 mm (3.75 ± 0.08, n=12). Biometric data
are given in Table 1. Body broadly oval and strongly convex. Cuticle
very strongly shining with green metallic lustre; pronotum black;
elytra with light blue or green metallic lustre; funicle of antennae
and tarsi dark brown; tibiae black.
Frontal stria of head (Fig. 1C) finely impressed, round anteriorly
but interrupted medially, its sides straight. Supraorbital stria
complete and well impressed. Disk of front densely covered with
coarse and deep punctures that are separated by their own diameter
and becoming sparser basally; a large deep puncture present at
medio-basal fourth. Epistoma more densely and finely punctate
than the medio-apical area of the frontal disk. Labrum broadly
and deeply depressed medially, its anterior margin emarginate
medially. Mentum (Fig. 2D) quadriangular, the anterior margin
roundly emarginate at middle.
Pronotal lateral sides (Fig. 1A) evenly convergent on basal five-sixths,
thence strongly convergent to apices. Apical angles round. Marginal
stria complete, its outer edge lightly carinate. Disk smooth medially
and densely covered with large, deep and oblong punctures along
lateral sides, the punctation on the lateral area expanded medio-anteriorly;
the punctures becoming finer and sparser medially and anteriorly;
basal area of disk with two or three rows of coarse punctures;
elsewhere finely and sparsely punctate, the punctures separated
by about four to ten times their diameter; area behind eyes strongly
depressed.
Epipleura of elytra (Fig. 1B) feebly convex; surface between epipleural
and elytral marginal striae covered sparsely with fine and oblong
punctures; inside elytral marginal stria sparsely covered with
fine punctures that are separated by four times their diameter.
Marginal epipleural stria complete, its ventral side subcariniform.
Marginal elytral stria complete, its apical end extending along
the posterior margin of elytron and united with the apical end
of sutural stria; its dorsal side strongly carinate. External
subhumeral stria confined to basal area. Internal subhumeral stria
absent. Oblique humeral stria well impressed and present on basal
third. First to third dorsal striae densely crenate with coarse
punctures, strongly impressed and present on basal third; second
a little longer than the first; first a little longer than the
third. Fourth dorsal stria present on basal half, it basal end
united with sutural stria as an arc. Fifth dorsal stria absent.
Sutural stria complete, deeply impressed and densely crenate.
Surface of elytra densely covered with large, round, and deep
punctures that are separated by their own to three times their
diameter, except on flanks, mediobasal half, and extreme apex
(outside apical stria); elsewhere sparsely and finely punctate,
the punctures separated by three to five times their diameter;
interval among the large punctures intermingled with fine punctures;
area between first and second dorsal striae densely with oblique
rugae.
Propygidium (Fig. 1D) short; disk densely covered with coarse
and shallow punctures that are separated by one-third to their
own diameter; on the basal half the punctures becoming finer.
Pygidium (Fig. 1D) rather sparsely covered with large, round and
shallow punctures that are about twice their diameter and becoming
finer apically; lateral margin feebly elevated; usually with narrow,
median longitudinal line on apical half which is impunctate. Interspace
among the punctures of propygidium and pygidium with altaceous
ground sculptures.
Anterior margin of median portion of prosternum (Fig. 2B) nearly
straight and shortly marginate on median third. Ventral disk of
keel depressed at apical third and slightly convex on basal two-thirds;
its disk finely covered with punctures that are separated by about
three times their diameter. Anterior marginal stria of keel strongly
impressed and complete, and its outer side strongly carinate.
Carinal striae divergent posteriorly and anteriorly and nearly
parallel on apical third; outer edge of stria distinctly carinate.
Descending lateral stria complete and its outer edge strongly
elevated, its apical end attaining to apical third of keel.
Anterior margin of mesosternum (Fig. 2B) slightly emarginate medially;
marginal stria interrupted medially and its outer side well carinate.
Disk sparsely covered with coarse punctures that are separated
by twice to five times their diameter. Meso-metasternal suture
distinctly impressed, complete and sparsely and coarsely crenate.
Intercoxal disk of metasternum (Fig. 2A) feebly convex, but strongly
depressed along longitudinal sutural line in male, while feebly
depressed in female; median area sparsely covered with fine punctures
that are separated by five to eight times their diameter; lateral
area along the lateral stria sparsely covered with coarse, round
and deep punctures, and with other fine ones intermingled; similarly
punctate in an apical transverse band, the punctures becoming
sparser and finer on median third; interspace among punctures
on lateral area and apical transverse band with finely impressed,
alutaceous micro-sculpture. Lateral metasternal stria strongly
carinate, extending obliquely and posteriorly, and present on
basal two-thirds. Lateral metasternal disk densely covered with
large, round, deep punctures that are separated by half to three
times their diameter, and with fine ones intermingled. Metepisternum
densely covered with large, deep, and longitudinal punctures,
and with fine punctures intermingled; interspace among these punctures
smooth and shining.
Intercoxal disk of 1st abdominal sternum covered with coarse punctures
along antero-lateral margins, which become sparser and finer medially;
lateral stria abbreviated at apical sixth and its outer edge subcariniform.
Protibia (Fig. 1E) with 10 to 12 spinules on outer margin, the
apical two and basal one small. Meso- (Fig. 1F) and metatibia
with 11 spinules on outer margin.
Male genitalia as shown in Fig. 3A-I; eighth sternite densely
with short setae; ratio in length of parameres to basal piece
about 4.0.
Female genitalia as shown in Fig. 3J; spermatheca globoid with
a deep invagination at each pole.
Specimens examined. [Proper] Nantou Hsien: Nanshanchi, Puli (4
exs., 12, 13, 14, 15/iv/1986), M. Ôhara; Ditto (1 ex., 6/vi;
22 exs., 9/vi; 1 ex., 17/vi/; 8 exs., viii/1972), Q. Du &
K. Masumoto; Ditto (1 ex., 30/v/1973), Q. Du; Shizuto, Puli (15
exs., 31/x/1987), K. Lah. Chiai Hsien: Abassen, Takesaki (= Chuchi)
(2 exs., 1/viii/38) (NSMT); Ditto (4 exs., 1/viii/1938), Y. Yano.
Kaohsiung Hsien: Liukuei (1 ex., 10/vii/1990), S. Osawa; Ditto
(1 ex., 37/v/1986), M. Ôhara. Pintung Hsien: Kenting
National Park (1 ex., 11/xi/1976), M. Kiuchi; Kuraru (= Kueitzuchiao),
(1 ex., 28/xii/1925), T. Kano (Kano collection, NSMT).
Remarks. S. optabilis commonly found under dead carcasses in Taiwan.
Mazur and Ohara (2004)
Specimens examined. Thailand: Poi Pui, Chiang Mai,
2.II.1988, Y. Manti, 15 exs.; Doi Suthep, Chiang Mai, 6.II.1989,
9.XI.1995, K. Masumoto, 2 exs.; Ban Huai Mo, 15.III.1992, Y. Okushima,
1 ex.; Doi Inthtanon, Chiang Mai, 14.III.1988, 2200 m. K. Masumoto,
5 exs.; Ban Phron Song, Chaiyaphunm Prov., 2.XII.1998, K. Masumoto,
3 exs.